Steps to reproduce: 
1. Add RadTreeView with few items 
2. Apply custom filtering 
3. In order to remove the custom filtering and return the default filtering, set the FilterPredicate to null. 
As a result, it is thrown an exception. 

Workaround: 
1. Save the default filter predicate before applying the custom filtering 
2. Clear filter descriptors and set the FilterPredicate property to stored default filter predicate

In the attachments can be found a sample application demonstrating the issue and the workaround